Anke Ninija Karabanov



+45 3862 2975


Senior Researcher, Associate Professor


University of Copenhagen, Integrative Physiology

Research Interests

I am interested in how the brain controls voluntary movemnts and how brain plasticity can optimally support skill learning. Methodologically I work with noninvasive brain stimulation, functional Magnetic Resonance Imaging (fMRI), Electroecephlography (EEG) and a wide gange of behavioral paradigms testing manual control and motor learning.
I am interested in how the brain contols skilled movemnts. I use neuroimaging and neurostimulation techniques to investigate the activity and connectvity patterns in the brain that support good motor control.

Currently, I work on brain state-dependnet exitability and connectivity patterns that can influence neuronal plasticity. The importance of frontoparietal connectivity for good sensori-motor integration and the effects dosage and placemnt of neuromodulatory TDCS on peripheral and central mechanisms.

Curriculum Vitae



Doctor of Philosophy (PhD) in Medical Sciences, Join Doctorate Program National Institutes of Health, USA and Karolinska Institute, Sweden


Bachelor of Cognitive Sciences (BcS), University of Osnabrück, Germany


2019 - Present

Associate Professor, Integrative Physiology, University of Copenhagen

2015 - Present

Senior Researcher, DRCMR

2011 - 2014

Postdoctoral Fellow, DRCMR

2010 - 2011

Postdoctoral Fellow, National Institute of Mental Health

Full CV


Video Pitch: Presentation

Kesselheim, J., Takemi, M., Karabanov, A. & Siebner, H. R.
Is there a causal link between afferent inhibition and oscillatory brain stimulation in the human sensorimotor cortex? 2018.

Karabanov, A. N., Ritterband-Rosenbaum, A., Christensen, M. S., Siebner, H. R. & Nielsen, J. B.
Modulation of fronto-parietal connections during the rubber hand illusion. European Journal of Neuroscience. 45, 7, p. 964-974, 2017.

Dubbioso, R., Raffin, E., Karabanov, A., Thielscher, A. & Siebner, H. R.
Centre-surround organization of fast sensorimotor integration in human motor hand area. NeuroImage. 158, p. 37-47, 2017.

Safeldt, M. G., Tomasevic, L., Karabanov, A., Siebner, H. R. & Madsen, K. H.
Towards Brain-State Dependent Transcranial Magnetic Stimulation: targeting the phase of oscillatory neocortical activity with singe-pulse TMS.

Safeldt, M. G., Karabanov, A., Tomasevic, L., Siebner, H. R. & Madsen, K. H.
Evaluation of phase estimator performances on simulated EEG for phase dependent TMS purposes. 2016.

Kesselheim, J., Takemi, M., Jønsson, M. G., Karabanov, A. & Siebner, H. R.
Exploiting I-wave rhythmicity to optimize the stimulus-response relationship for TMS of M1. 1 p., 2016.

Kesselheim, J., Takemi, M., Jønsson, M. G., Karabanov, A. & Siebner, H. R.
Exploiting I-Wave Rhythmicity To Optimizing The Stimulus-Response Relationship For TMS Of M1. 2016.

Karabanov, A., Irmen, F., Bisgaard, T. & Siebner, H. R.
Functional Plasticity During De Novo Acquisition of Laparoscopic Surgical Skills. 2016.

Dubbioso, R., Raffin, E. E., Karabanov, A., Haahr Nielsen, S., Thielscher, A. & Siebner, H. R.

Karabanov, A.
Hjerneforskerens gode råd: Sådan afværger du vinterdepressionen. Søndagsavisen A/S. 2016.

Karabanov, A.
Sådan slipper du for vinter-depressionen. Hvidovre Lokalhistorie. 2016.

Karabanov, A., Lohse, A. & Siebner, H. R.
Hjernestimulation og Den forudsigende hjerne. Den forudsigende hjerne. Hjerneforum, p. 46-52, 2016.

Karabanov, A., Thielscher, A. & Siebner, H. R.
Transcranial brain stimulation: closing the loop between brain and stimulation. Current Opinion in Neurology. 29, 4, p. 397-404, 2016.

Bergmann, T. O., Karabanov, A., Hartwigsen, G., Thielscher, A. & Siebner, H. R.
Combining non-invasive transcranial brain stimulation with neuroimaging and electrophysiology: Current approaches and future perspectives.
NeuroImage. 140, p. 4-19, 2016.

Hartwigsen G, Bergmann TO, Herz DM, Angstmann S, Karabanov AN, Raffin E, Thielscher A, Siebner H. (accepted) Modeling the effects of non-invasive transcranial brain stimulation at the biophysical, network and cognitive level. Progressive Brain Research.

Karabanov AN, Raffin E, Siebner HR (2015) The Resting Motor Threshold - Restless or Resting?  Brain Stimulation, Epub. Ahead of print

Karabanov AN, Ziemann U, Hamada M, George MS, Quartarone A, Classen J, Massimini M, Rothwell J, Siebner HR (2015) Probing Homeostatic Plasticity of Human Cortex With Non-invasive Transcranial Brain Stimulation. Vol. 8(3) p. 442-54

Karabanov AN, Paine R, Chao CC, Schulze K, Scott B, Hallett M, Mishkin M (2015) Participation of the classical speech areas in auditory long-term memory. PLoS One. Vol. 27;10(3) e:0119472

Chao CC, Karabanov AN, Paine R, Carolina de Campos A, Kukke SN, Wu T, Wang H, Hallett M. (2015) Induction of Motor Associative Plasticity in the Posterior Parietal Cortex-Primary Motor Network. Cerebal Cortex, Vol. 25(2) p.365-73

Karabanov AN, Siebner, HR (2014) Expanding the electrotherapeutic toolkit, Acta Neuropsychiatrica Vol.26(5) p.261-3

Karabanov AN, Ritterband-Rosenbaum A, Christensen MS, Nielsen (2014) 10 Hz rTMS over right parietal cortex alters sense of agency during self-generated movements. Frontiers Human Neuroscience, Vol. 25(8) e:417

Karabanov AN, Siebner HR (2014) Unravelling homeostatic interactions in inhibitory and excitatory networks in human motor cortex. Journal of Physiology, Vol. 15(590) p.5557-8

Karabanov AN, Chao CC, Paine R, Hallett M. (2013). Mapping different intra-hemispheric parietal-motor networks using twin Coil TMS.  Brain Stimulation Vol.6(3) p.384-9

Houdayer E, Beck S, Karabanov AN, Poston B, Hallett M. 2012. The differential modulation of the ventral premotor-motor interaction during movement initiation is deficient in patients with focal hand dystonia. European Journal of Neuroscience, Vol. 5(3) p.478-85

Karabanov AN, Jin SH, Joutsen A, Poston B, Aizen J, Ellenstein A, Hallett M. (2012) Timing-dependent modulation of the posterior parietal cortex-primary motor cortex pathway by sensorimotor training. Journal of Neurophysiology, Vol. 07(11) p. 3190-9

Forsman LJ, de Manzano O, Karabanov AN, Madison G, Ullén F, Differences in regional brain volume related to the extraversion-introversion dimension – a voxel based morphometry study. Neuroscience Research 72(1):59-67

Karabanov AN, Cervenka S, DeManzano O, Forssberg H, Farde L, Ullen F. 2010. Dopamine D2 receptor density in the limbic striatum is related to implicit but not explicit movement sequence learning. Proc Nat Ac Sci USA. Vol 20;107(16) p. 7574-9.

de Manzano O, Cervenka S, Karabanov AN, Farde L, Ullén F (2010) Thinking utside a less intact box: thalamic dopamine D2 receptor densities are negatively related to psychometric creativity in healthy individuals. PLoS One. Vol. 17;5(5) e:10670

Karabanov AN, Blom O, Forsman L, Ullén F. (2009). The dorsal auditory pathway is involved in performance of both visual and auditory rhythms. Neuroimage. Vol. 15;44(2) p.480-8

Karabanov AN, Ullén F, Implicit and explicit learning of temporal sequences studied with the process dissociation procedure. Journal of Neurophysiology, Vol. 100(2) p.733-9

Madison G, Forsman L, Blom O, Karabanov AN, Ullen F. (2009) Correlations between intelligence and components of serial timing. Intelligence. Vol. 37 p.68-75.

Ullén F, Forsman L, Blom O, Karabanov AN, Madison G. Intelligence and variability in a simple tapping task share neural substrates in the prefrontal white matter. Journal of Neuroscience. Vol. 28(16) p.4238-43